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Chuck Shurley is the pseudonym used by God when he was living as an author who wrote the low selling Supernatural book series, which he wrote under the nom de plume Carver Edlund.
Initially Chuck claimed to the Winchesters that he foresaw events that concerned Dean and Sam, with Castiel revealing that Chuck was a prophet. He tells the Winchesters that he goes through some kind of a "process" when he has visions -- a severe headache occurs, and then he takes aspirin and drinks until he passes out. While unconscious, he says he sees and interprets prophecy.
After the Darkness was released, Chuck returned to reveal to Metatron that He is in fact God in human disguise. He told Metatron that he had taken on human form so he could "hide in plain sight", as he likes having front row seats.
Chuck reveals himself as God to Metatron and asks for help in writing a book and Metatron asks Chuck many questions and Chuck finally decides to be himself as God instead of just being Chuck. He appears in every other scene with Metatron. The scenes switch from Sam and Dean and everyone in the town they are in and then it goes to Chuck and Metatron and then keeps switching between them. Chuck reveals himself as God to the Winchesters at the end of the episode through Dean's Amulet.